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Simpson’s Diversity Index Calculator (Free) | StatsUnlock

Simpson's Diversity Index Calculator (Free) | StatsUnlock

Simpson's Diversity Index Calculator

A free online tool to measure species diversity with Simpson's index (D, 1-D, and inverse Simpson) from your survey counts, with instant charts and report-ready results.

BiodiversityDiversity IndexDominanceEcologyFreeOnline

Last updated: 15 September 2026 · Built for ecologists and wildlife researchers by StatsUnlock

⚡ 0. Quick Answer

The Simpson's diversity index calculator turns species counts into a diversity value between 0 and 1. Enter counts such as 45, 30, 15, 7, 3 and the tool returns dominance D = 0.321 from D = Σ pi², diversity 1 - D = 0.679, and reciprocal 1/D = 3.12. Higher 1 - D means greater diversity.

🌿 Key Takeaways

  • The Simpson's diversity index calculator turns your species counts into a diversity number based on how much the community is dominated by common species.
  • You only need the count of individuals per species. Paste them comma separated, upload a CSV file, or type them in.
  • It returns three linked values: Simpson's dominance (D), Simpson's index of diversity (1-D), and inverse Simpson (1/D).
  • A higher 1-D means greater diversity and less dominance; a value near 0 means one or two species dominate the sample.
  • Typical field uses: camera trapping, point counts, vegetation plots, transects, and reef fish surveys.

🌱 1. What Is Simpson's Diversity Index?

Simpson's diversity index answers one plain question: if you pick two individuals at random from your sample, what is the chance they belong to the same species? That chance is dominance, written D. Subtract it from 1 and you get Simpson's index of diversity, 1 - D, the chance the two individuals are different species.

The number comes from proportions. Divide each species count by the total to get a proportion p, square each one, and add them up. That sum is D. Because D is a probability, it always sits between 0 and 1, which makes Simpson's index easier to explain to a non-specialist than the Shannon index.

Ecologists use it because it is dominated by the common species. A community with one species holding most individuals gets a high D and a low 1 - D, no matter how many rare species are hiding in the tail. That is exactly what you want when the question is about disturbance, invasion, or one species taking over.

Even community One species dominates two random picks rarely match two random picks usually match D = 0.20, 1 - D = 0.80 D = 0.66, 1 - D = 0.34 D is the chance of a match. 1 - D is the chance of a difference.
Simpson's index in one picture. Both samples hold ten individuals; only the balance between species changes.

1.1 The three numbers people call "Simpson's index"

NameSymbolRangeReads as
Simpson's dominanceD0 to 1Higher means one species dominates
Simpson's index of diversity1 - D0 to 1Higher means more diverse. This is the one most papers report.
Simpson's reciprocal index1/D1 to SEffective number of common species

What to conclude: always say which form you are reporting. "Simpson's index = 0.32" and "Simpson's index = 0.68" can describe the very same community, which is the single most common source of confusion in the literature.

1.2 What the value means

1 - D rangeReadingTypical setting
0.00 to 0.30Very low ConcernMonoculture, one invasive species, heavily polluted water
0.30 to 0.50Low ConcernDegraded fragment, intensive farmland, urban core
0.50 to 0.70Moderate WatchSecondary forest, managed grassland, recovering site
0.70 to 0.85High HealthyMature temperate forest, healthy wetland, good reef transect
Above 0.85Very high HealthyLowland tropical forest, large insect trap samples

What to conclude: these bands are a first reading only. Compare your value against a reference site sampled with the same method, the same effort, and the same season.

📥 2. Set Up Your Data

0 values entered

Supports .csv and .txt, headers detected automatically.
SpeciesCount

🔎 4. Interpretation of Results, In Detail

▶ Run the analysis above to generate a detailed, plain-English interpretation of your Simpson's diversity result.

✍️ 5. How to Write Your Results in Research

▶ Run the analysis above to auto-fill all six examples with your results.

🪧 5.1 Research Poster Panel

A print-ready poster layout that updates with your result. Copy the text or use the visual scaffold for A0 / A1 conference posters.

▶ Run the analysis above to build your poster panel.

🧮 6. Formulas Used

Every number this Simpson's diversity index calculator reports comes from one of the seven formulas below. Card 1 is dominance, and each later card moves one step outward.

1

Simpson's dominance (the infinite population form)

D = Σi=1S pi²
  • D dominance, the chance two random individuals are the same species, unitless
  • pi proportion of individuals in species i, unitless
  • S number of species in the sample, a count
  • Range: 1/S to 1. D = 1 means a single species.
  • Example: counts 45, 30, 15, 7, 3 give D = 0.321.
2

Simpson's index of diversity

1 - D = 1 - Σ pi²
  • 1 - D the chance two random individuals are different species, 0 to 1
  • Rule: this is the form most journals expect. Say so explicitly in your methods.
  • Example: 1 - 0.321 = 0.679.
  • Note: sometimes written as the Gini-Simpson index.
3

Simpson's reciprocal index

1/D = 1 / Σ pi²
  • 1/D effective number of common species, in species units
  • Range: 1 to S. Equals S only when every species is equally common.
  • Use for: honest ratios. 1/D of 8 really is twice 1/D of 4.
  • Example: 1 / 0.321 = 3.12 effective species out of 5 observed.
4

The finite sample form (use this for real field data)

D = Σ ni(ni - 1) / [ N(N - 1) ]
  • ni count of individuals of species i
  • N total individuals across all species
  • Requires: whole-number counts, never percentages.
  • Example: 3108 / 9900 = 0.314, slightly below the 0.321 the proportion form gives.
  • Rule: the two forms converge as N grows. Below about N = 100 the difference matters.
5

Species proportion

pi = ni / N
  • Rule: all proportions must add to 1.00.
  • Note: unlike the Shannon index, a species with a count of 0 causes no error here. It simply contributes nothing.
  • Use for: the input to cards 1, 2 and 3.
6

Simpson's evenness

E1/D = (1/D) / S
  • E1/D evenness, unitless, 0 to 1
  • Range: 1.0 is perfectly even, values under 0.5 mean strong dominance.
  • Example: 3.12 / 5 = 0.623.
  • Note: this is the Simpson equivalent of Pielou's J'.
7

Berger-Parker dominance, the quick screen

d = nmax / N
  • nmax count of the single commonest species
  • Range: 1/S to 1.
  • Example: 45 / 100 = 0.45, so the top species holds 45 percent of individuals.
  • Use for: a one-second check on whether dominance is driving your Simpson value.

🧭 7. How to Use This Calculator

Ten steps from a field sheet to a publication-ready Simpson's diversity result.

  1. Collect standardised counts. Record the number of individuals for each species using the same effort at every site.
  2. Pick an input tab. Use Paste or Type for a quick run, Upload CSV for a spreadsheet, or Manual Table to key in rows.
  3. Enter species names and counts. In Column Entry mode, put the species name on the left and the count on the right.
  4. Or load a sample dataset. Choose one of the built-in field datasets to see how the tool behaves before you use your own data.
  5. Name your study area. Type the site name into the Study Area box so it appears in every report template and in the poster panel.
  6. Decide which form you will report. 1 - D is the usual choice. Note it in your methods so nobody misreads your number.
  7. Set the decimal places. Three decimals is normal for Simpson's index in ecology papers.
  8. Click Run Analysis. The tool returns D, 1 - D, 1/D, richness, evenness, charts, and a results table.
  9. Read the interpretation. It explains your number in plain English and flags any sampling problems.
  10. Copy or download your result. Use the report templates, the poster panel, Download Doc, or Download PDF for your thesis or paper.

📗 8. How to Calculate Simpson's Diversity Index in Excel

Excel, step by step, with the same worked example used everywhere on this page

The worked example below uses five species with counts 45, 30, 15, 7, 3 (N = 100). Excel, R, Python and this calculator all return D = 0.321, 1 - D = 0.679 and 1/D = 3.12 for it.

8.1 The Excel functions you need

FunctionWhat it doesExample
SUMAdds your counts to get N=SUM(B2:B6)
SUMSQSquares every value and adds them, giving D in one cell=SUMSQ(C2:C6)
SUMPRODUCTBuilds the finite sample form without a helper column=SUMPRODUCT(B2:B6,B2:B6-1)/(B7*(B7-1))
COUNTIFCounts species with a count above zero, giving S=COUNTIF(B2:B6,">0")
MAXFinds the commonest species for Berger-Parker=MAX(B2:B6)/B7
ROUNDDisplays three decimals without losing precision upstream=ROUND(D7,3)

What to conclude: SUMSQ is the shortcut nobody teaches. It replaces an entire squared-proportion column with one cell.

8.2 Step by step in the sheet

Step 1. Type your data and total the counts

=SUM(B2:B6)
simpson.xlsx - ExcelB7fx=SUM(B2:B6)ABCDESpeciesCountp = n/Np^2n(n-1)12Bulbul453Babbler304Sunbird155Barbet76Drongo37Total N100

Species names go in column A, counts in column B. Cell B7 holds =SUM(B2:B6), which returns N = 100.

Step 2. Turn each count into a proportion

=B2/$B$7fill down to C6
simpson.xlsx - ExcelC2fx=B2/$B$7ABCDESpeciesCountp = n/Np^2n(n-1)12Bulbul450.453Babbler300.304Sunbird150.155Barbet70.076Drongo30.03

C2 holds =B2/$B$7 and returns 0.45. The dollar signs lock the total so the formula fills down correctly to C6. Column C now adds to 1.00.

Step 3. Square every proportion

=C2^2fill down to D6
simpson.xlsx - ExcelD2fx=C2^2ABCDESpeciesCountp = n/Np^2n(n-1)12Bulbul450.450.20253Babbler300.300.09004Sunbird150.150.02255Barbet70.070.00496Drongo30.030.0009

D2 holds =C2^2 and returns 0.2025. Squaring is what makes Simpson's index lean on the common species: 0.45 squared is 0.2025, but 0.03 squared is only 0.0009.

Step 4. Add the squares and read all three forms

=SUM(D2:D6)=SUMSQ(C2:C6)=1-D7=1/D7
simpson.xlsx - ExcelD7fx=SUM(D2:D6)ABCDESpeciesCountp = n/Np^2n(n-1)12Bulbul450.450.202519803Babbler300.300.09008704Sunbird150.150.02252105Barbet70.070.0049426Drongo30.030.000967D (dominance)0.3208

D7 holds =SUM(D2:D6) and returns D = 0.3208. Diversity is =1-D7 = 0.6792 and the reciprocal is =1/D7 = 3.117. Column E shows =B2*(B2-1) for the finite sample version.

Tip: the whole index fits in one cell. =1-SUMSQ(B2:B6)/SUM(B2:B6)^2 gives 1 - D straight from the raw counts, with no proportion column at all.
Warning: Excel will happily calculate Simpson's index from percentages, and the answer will be wrong by a factor of 10,000. The finite sample form needs whole-number counts. Check that B7 reads your true total, not 100 percent.

8.3 The awkward cases

CaseWhat breaksFix
A species with a count of 0Nothing breaks, but it inflates your species count SDelete the row, or use =COUNTIF(B2:B6,">0") for S
Only one species present1/D returns 1 and 1 - D returns 0That is correct. A single species has zero diversity.
Counts entered as percentagesThe finite sample form returns a meaningless valueGo back to the raw counts, or use the proportion form only
Text mixed into the count columnSUM ignores it silently, so N is wrongCheck N by eye against =COUNT(B2:B6)
Duplicate species rowsThe same species is split in two, lowering DUse a PivotTable to sum counts by species name first

What to conclude: Simpson's index is more forgiving than Shannon's because zeros do not throw errors. That makes silent mistakes easier to miss, so check N every time.

8.4 Why answers differ between Excel, R and Python

The usual cause is not software, it is which form was reported. R's vegan returns 1 - D by default when you ask for "simpson", and "invsimpson" for 1/D. Python's skbio returns 1 - D as well. If your Excel sheet stops at =SUMSQ, you are holding D, not 1 - D, and the two look nothing alike. The second cause is the finite sample correction: this page's worked example gives 0.3208 with proportions and 0.3139 with the n(n-1) form. Both are correct; state which you used. Rounding proportions to two decimals before squaring is the third cause, and it can shift the third decimal.

8.5 Charting the result in Excel

  1. Relative abundance pie. Select columns A and C, then Insert, Pie chart. Simpson's index is a dominance measure, so the pie is the honest picture of it.
  2. Contribution bar chart. Chart column D. The tallest bar is the species driving your D value, which is usually the sentence a reviewer wants.
  3. Site comparison bars. One 1 - D value per site in a column, then a clustered bar chart with the axis fixed from 0 to 1.
  4. Label the axis properly. Write "Simpson's 1 - D" on the value axis, never just "Simpson's index".

8.6 Excel error messages and what they mean

MessageCauseFix
#DIV/0!N is zero, or D is zero in the 1/D cellCheck the SUM range covers every data row
#VALUE!Text inside a numeric rangeClear notes such as "none seen" from the count column
#NUM!A negative count reached the squaring stepCounts cannot be negative. Re-check data entry.
Result is exactly 1You calculated D for a single-species sampleCorrect, but report 1 - D = 0 rather than D = 1
Result is above 1Percentages were used with the finite sample formSwitch to raw counts
1/D is smaller than 1D was entered above 1Impossible value. Recheck the proportion column adds to 1.00.

What to conclude: any Simpson value outside 0 to 1 is a data problem, not an ecological finding.

📘 9. How to Calculate Simpson's Diversity Index in R

R, one complete script you can run top to bottom

9.1 The complete script

Copy this into RStudio and press Run. It needs no packages. Change only the counts line. It prints all three Simpson forms and saves one 300 dpi figure.

# ------------------------------------------------------------------
# Simpson's Diversity Index in R
# StatsUnlock.com - free ecology and wildlife analysis tools
# ------------------------------------------------------------------
set.seed(42)

# 1. YOUR DATA -- change only this line
counts <- c(45, 30, 15, 7, 3)
species <- c("Bulbul", "Babbler", "Sunbird", "Barbet", "Drongo")

# Reading from a file instead? Uncomment the two lines below.
# dat <- read.csv("my_survey.csv")          # columns: species, count
# counts <- dat$count; species <- dat$species

# 2. CLEAN THE DATA
counts[is.na(counts)] <- 0        # missing values become zero
keep <- counts > 0                # zero counts add nothing, drop them
counts <- counts[keep]
species <- species[keep]

# 3. CALCULATE
N <- sum(counts)                          # total individuals
S <- length(counts)                       # species richness
p <- counts / N                           # proportions
D <- sum(p^2)                             # Simpson's dominance
div <- 1 - D                              # Simpson's index of diversity
inv <- 1 / D                              # Simpson's reciprocal index
D_finite <- sum(counts * (counts - 1)) / (N * (N - 1))   # finite sample form
even <- inv / S                           # Simpson's evenness
bp <- max(counts) / N                     # Berger-Parker dominance

# Same answers using the vegan package:
# install.packages("vegan")
# library(vegan)
# vegan::diversity(counts, index = "simpson")      # returns 1 - D
# vegan::diversity(counts, index = "invsimpson")   # returns 1 / D

# 4. PRINT A PLAIN-ENGLISH VERDICT
verdict <- if (div >= 0.70) "high diversity" else if (div >= 0.50) "moderate diversity" else "low diversity"
cat("Dominance D             :", round(D, 3), "\n")
cat("Simpson 1 - D           :", round(div, 3), "\n")
cat("Reciprocal 1/D          :", round(inv, 3), "\n")
cat("Finite sample D         :", round(D_finite, 3), "\n")
cat("Species richness S      :", S, "\n")
cat("Total individuals N     :", N, "\n")
cat("Simpson evenness        :", round(even, 3), "\n")
cat("Berger-Parker d         :", round(bp, 3), "\n")
cat("Verdict                 :", verdict, "\n")

# 5. SAVE ONE 300 dpi FIGURE
png("simpson_contribution.png", width = 1800, height = 1200, res = 300)
ord <- order(p^2, decreasing = TRUE)
barplot(p[ord]^2,
        names.arg = species[ord],
        col = "#3aab6d", border = NA, las = 2,
        ylab = "Contribution to D (p squared)",
        main = paste0("What drives D, 1 - D = ", round(div, 3)))
dev.off()
cat("Figure saved as simpson_contribution.png\n")

9.2 What each line does

LineWhat it does
set.seed(42)Fixes anything random so your figure is reproducible.
counts <- c(45, 30, ...)Your species counts. This is the only line most users change.
keep <- counts > 0Drops zero counts. They add nothing to D, but they inflate S.
p <- counts / NConverts counts to proportions that add to 1.00.
D <- sum(p^2)Simpson's dominance. The core of the whole index.
div <- 1 - DSimpson's index of diversity, the form most papers report.
inv <- 1 / DSimpson's reciprocal index, the effective number of common species.
D_finite <- sum(counts * (counts - 1)) / (N * (N - 1))The finite sample form. Use it when N is small.
diversity(x, "simpson")The vegan one-liner. It returns 1 - D, not D. This trips people up constantly.
diversity(x, "invsimpson")The vegan call for 1/D.
dev.off()Closes the device and writes the file. Forget this and the PNG stays empty.

9.3 What the figure shows

The script saves a contribution chart: each species plotted by its squared proportion, sorted largest first. This is the most useful figure for Simpson's index, because it names the species responsible for your D value. In the worked example the first bar alone (0.2025) is nearly two thirds of D = 0.3208, which tells the whole story in one glance.

9.4 Problems and fixes

ProblemCauseFix
vegan gives 0.679 but my sheet gives 0.321vegan returns 1 - D, your sheet returned DBoth are right. Decide which form you report and be consistent.
Error: could not find function "diversity"vegan is not loadedRun install.packages("vegan") then library(vegan)
1/D returns InfD is zero, which needs an empty sampleCheck that counts holds at least one positive value
Value changes when I add rare speciesExpected. Simpson's index barely moves for rare species.If rare species matter, report richness or the Shannon index too
Empty PNG filedev.off() was never calledRun dev.off() once, then re-run the plotting block
Finite form differs from veganvegan uses the proportion formExpected below N = 100. State which form you used.

9.5 One-liners worth keeping

CodeReturns
vegan::diversity(x, "simpson")1 - D from a count vector or a site-by-species matrix
vegan::diversity(x, "invsimpson")1/D, the reciprocal index
1 - vegan::diversity(x, "simpson")D, the dominance form
vegan::specnumber(x)Species richness S
max(x)/sum(x)Berger-Parker dominance
iNEXT::iNEXT(x, q = 2)Rarefaction and extrapolation for Hill number q = 2, which equals 1/D

🐍 10. How to Calculate Simpson's Diversity Index in Python

Python, one complete script you can run top to bottom

Packages: numpy does the arithmetic and matplotlib draws and saves the figure. Install them once with pip install numpy matplotlib. scikit-bio is optional and gives a one-line check. pandas is only needed if you read a CSV file. SciPy has no direct Simpson function, which is why the manual line matters here.

10.1 The complete script

Save this as simpson.py and run it. Change only the counts line. It prints all three Simpson forms and saves one 300 dpi figure.

# ------------------------------------------------------------------
# Simpson's Diversity Index in Python
# StatsUnlock.com - free ecology and wildlife analysis tools
# ------------------------------------------------------------------
import numpy as np
import matplotlib
matplotlib.use("Agg")
import matplotlib.pyplot as plt

np.random.seed(42)

# 1. YOUR DATA -- change only this line
counts = np.array([45, 30, 15, 7, 3], dtype=float)
species = ["Bulbul", "Babbler", "Sunbird", "Barbet", "Drongo"]

# Reading from a file instead? Uncomment the three lines below.
# import pandas as pd
# dat = pd.read_csv("my_survey.csv")          # columns: species, count
# counts, species = dat["count"].to_numpy(float), dat["species"].tolist()

# 2. CLEAN THE DATA
counts = np.nan_to_num(counts, nan=0.0)       # missing values become zero
keep = counts > 0                             # zero counts add nothing
species = [s for s, k in zip(species, keep) if k]
counts = counts[keep]

# 3. CALCULATE
N = counts.sum()                              # total individuals
S = counts.size                               # species richness
p = counts / N                                # proportions
D = float(np.sum(p ** 2))                     # Simpson's dominance
div = 1 - D                                   # Simpson's index of diversity
inv = 1 / D                                   # Simpson's reciprocal index
D_finite = float(np.sum(counts * (counts - 1)) / (N * (N - 1)))   # finite sample
even = inv / S                                # Simpson's evenness
bp = float(counts.max() / N)                  # Berger-Parker dominance

# Same answer using scikit-bio:
# from skbio.diversity.alpha import simpson
# div = simpson(counts.astype(int))    # returns 1 - D

# 4. PRINT A PLAIN-ENGLISH VERDICT
verdict = "high diversity" if div >= 0.70 else "moderate diversity" if div >= 0.50 else "low diversity"
print(f"Dominance D             : {D:.3f}")
print(f"Simpson 1 - D           : {div:.3f}")
print(f"Reciprocal 1/D          : {inv:.3f}")
print(f"Finite sample D         : {D_finite:.3f}")
print(f"Species richness S      : {S}")
print(f"Total individuals N     : {N:.0f}")
print(f"Simpson evenness        : {even:.3f}")
print(f"Berger-Parker d         : {bp:.3f}")
print(f"Verdict                 : {verdict}")

# 5. SAVE ONE 300 dpi FIGURE
contrib = p ** 2
order = np.argsort(contrib)[::-1]
fig, ax = plt.subplots(figsize=(6, 4))
ax.bar([species[i] for i in order], contrib[order], color="#3aab6d")
ax.set_ylabel("Contribution to D (p squared)")
ax.set_title(f"What drives D, 1 - D = {div:.3f}")
plt.xticks(rotation=45, ha="right")
plt.tight_layout()
plt.savefig("simpson_contribution.png", dpi=300)
print("Figure saved as simpson_contribution.png")

10.2 What each line does

LineWhat it does
matplotlib.use("Agg")Draws without a screen, so the script runs on a server or in a notebook.
counts = np.array([...])Your species counts. The only line most users change.
np.nan_to_num(counts)Turns missing values into zero so the script does not stop.
keep = counts > 0Drops zero counts, which add nothing to D but inflate S.
p = counts / NConverts counts to proportions that add to 1.00.
D = np.sum(p ** 2)Simpson's dominance.
div = 1 - DSimpson's index of diversity, the form most papers report.
inv = 1 / DSimpson's reciprocal index.
D_finite = ...(counts - 1)...The finite sample form, for small N.
skbio ... simpson(counts)The scikit-bio one-liner. It returns 1 - D and needs integer counts.
plt.savefig(..., dpi=300)Writes the 300 dpi figure most journals ask for.

10.3 What the figure shows

The saved PNG plots each species by its contribution to D, sorted largest first, identical in meaning to the R figure. If one bar towers over the rest, dominance is driving your result and 1 - D will be low. For a relative abundance pie instead, swap ax.bar for ax.pie(p, labels=species).

10.4 Problems and fixes

ProblemCauseFix
ImportError: cannot import simpsonscikit-bio is not installed, or the name changed between versionsUse the manual NumPy line, which needs no extra package
scipy.stats has no simpson for diversityscipy.integrate.simpson is numerical integration, not diversityDo not use it. Calculate D with NumPy directly.
Result is 0.321 not 0.679You printed D rather than 1 - DReport div, and say which form in your methods
ZeroDivisionError on 1/DEvery count was filtered outCheck the array is not empty after the keep filter
Integer division gives zerosCounts stored as int in old Python codeKeep dtype=float in the array line
Figure window never opensThe Agg backend has no screen outputThat is expected. Open the saved PNG file.

10.5 One-liners worth keeping

CodeReturns
1 - np.sum((c/c.sum())**2)Simpson's 1 - D, no packages beyond NumPy
1 / np.sum((c/c.sum())**2)Simpson's reciprocal index 1/D
np.sum(c*(c-1))/(c.sum()*(c.sum()-1))The finite sample dominance D
skbio.diversity.alpha.simpson(c)1 - D from scikit-bio, integer counts only
c.max()/c.sum()Berger-Parker dominance
(c > 0).sum()Species richness S

📋 11. Reference Tables

Six lookup tables for reading, converting, and sanity-checking a Simpson's diversity index value.

Table 11.1 Published 1 - D values by habitat type

HabitatTaxonTypical 1 - DTypical S
Lowland tropical rainforestBirds0.90 to 0.9640 to 120
Dry deciduous forestBirds0.82 to 0.9225 to 50
Temperate woodlandBirds0.78 to 0.8815 to 35
Grassland, managedPlants0.65 to 0.8510 to 30
Wetland, freshwaterWaterbirds0.72 to 0.8812 to 40
Coral reef transectFish0.85 to 0.9430 to 80
Camera trap gridLarge mammals0.60 to 0.826 to 20
Urban parkBirds0.55 to 0.808 to 20
Intensive croplandArthropods0.45 to 0.755 to 20
Polluted streamMacroinvertebrates0.15 to 0.553 to 10

What to conclude: Simpson values cluster high and compress at the top. A jump from 0.90 to 0.94 is a bigger ecological change than it looks, which is why the reciprocal form is easier to compare.

Table 11.2 Converting between the three Simpson forms

D (dominance)1 - D (diversity)1/D (reciprocal)Reading
1.0000.0001.00One species only
0.8000.2001.25Extreme dominance
0.5000.5002.00Strong dominance
0.3210.6793.12The worked example on this page
0.2500.7504.00Moderate
0.2000.8005.00High
0.1000.90010.00Very high
0.0500.95020.00Exceptional
0.0200.98050.00Large, very even sample

What to conclude: 1 - D changes very little between 0.95 and 0.98, while 1/D doubles from 20 to 50. For comparing sites, report 1/D as well.

Table 11.3 Maximum possible diversity for a given richness

Species richness SMinimum D (all equal)Maximum 1 - DMaximum 1/D
20.5000.5002
50.2000.8005
100.1000.90010
200.0500.95020
500.0200.98050
1000.0100.990100

What to conclude: 1/D can never exceed S. If your reported 1/D is larger than your species count, you have a calculation error.

Table 11.4 Reading Simpson's evenness

E = (1/D) / SReadingWhat it usually means
0.90 to 1.00Very even HealthyNo species dominates. Common in stable systems.
0.75 to 0.90Even HealthyA normal, balanced community.
0.60 to 0.75Moderate WatchTwo or three species carry most individuals.
0.40 to 0.60Uneven WatchClear dominance. Check for disturbance or an aggregation.
Below 0.40Very uneven ConcernOne species takes over. Often invasion or pollution.

What to conclude: the worked example scores 0.623, moderate. Two species out of five hold three quarters of the individuals.

Table 11.5 Minimum sampling effort by survey type

Survey typeMinimum effort for a usable 1 - DPreferred effort
Bird point counts10 points, 30 individuals30 points across 3 visits
Line transects2 km walked10 km, repeated in 2 seasons
Camera trapping300 trap nights1,000 to 2,000 trap nights
Vegetation quadrats10 quadrats of 1 m²30 quadrats, stratified by habitat
Pitfall traps20 trap days100 trap days across 5 stations
Reef fish belt transects4 transects of 50 m12 transects across 3 depths
Light trap insects3 trap nights12 trap nights across the season

What to conclude: Simpson's index needs less effort than the Shannon index to stabilise, because it depends on the common species. That is one of its real advantages.

Table 11.6 Which diversity index to report

IndexWeightsRangeBest used when
Simpson's 1 - DCommon species0 to 1Dominance is the story, or the sample is small
Simpson's 1/DCommon species1 to SYou need a comparable ratio between sites
Shannon Wiener H'Rare species0 to ~4.6You want richness and evenness balanced in one number
Species richness SAll equally1 upwardPresence lists only, no reliable counts
Pielou's J'Evenness only0 to 1Comparing communities with different richness
Berger-ParkerThe single commonest species0 to 1Quick screening for one dominant species
Hill numbersTunable by q1 upwardYou need a true diversity ratio. 1/D is Hill q = 2.

What to conclude: most reviewers expect Simpson plus one companion index. Simpson with Shannon is the safest pair, because one leans on common species and the other on rare ones.

📊 12. Example Results

Scenario 1: High-diversity tropical forest

A bird survey records 28 species and 412 individuals with fairly even counts. Simpson's 1-D works out to about 0.95. This high value means two random birds are almost always different species, typical of an intact community.

Scenario 2: Disturbed, degraded site

A logged fragment holds 6 species and 200 individuals, but one species makes up 150 of them. Simpson's 1-D drops to about 0.42. The low value reflects strong dominance and reduced ecological integrity.

Scenario 3: Same richness, different evenness

Two plots each have 5 species. Plot A counts 20,20,20,20,20 (1-D = 0.80); Plot B counts 80,5,5,5,5 (1-D = 0.35). Same richness, very different diversity, Simpson's index reacts strongly to the dominant species.

Scenario 4: Before vs after restoration

A grassland plot rises from 1-D = 0.55 before management to 1-D = 0.78 three years after invasive removal, showing recovering evenness as native species return.

Scenario 5: Camera-trap mammal survey

Across 1,400 trap nights, 8 mammal species are detected with counts 40,35,30,25,20,15,10,5. Simpson's 1-D = 0.83, a fairly high value for a mid-sized mammal community in a wildlife corridor.

Scenario 6: Edge case / caution

Only 3 individuals across 3 species gives 1-D = 1.00, but the sample is far too small to trust. Always collect at least 30 individuals before reporting a Simpson's value.

Scenario 7: Where 1 - D flattens and 1/D does not

Site X scores 1 - D = 0.95 and Site Y scores 1 - D = 0.98. The gap looks trivial. In reciprocal form they are 20.0 and 50.0 effective species, a two and a half times difference. This is why rich communities should always be compared with 1/D.

Scenario 8: The worked example used in Sections 8, 9 and 10

Five species with counts 45, 30, 15, 7, 3 give N = 100, S = 5, D = 0.321, 1 - D = 0.679, 1/D = 3.12, Simpson evenness 0.623 and Berger-Parker 0.45. The finite sample form gives D = 0.314. Excel, R, Python and this calculator all agree, so use this dataset to check your own setup.

🥾 13. How to Collect Raw Data in the Field

Simpson's index is only as good as the counts you feed it. Follow these ready-to-use steps in the field to collect clean, comparable species abundance data. Print this list and take it with you.

start end Line Transect Walk the line, record perpendicular distance
Fig 1. Line transect. Walk a fixed straight line at steady pace and count every individual seen or heard, noting its distance from the line.
observer radius r Point Count Stand still, count within a fixed radius
Fig 2. Point count. Stand at a fixed station for a set time (e.g. 10 min) and count all individuals within a fixed radius; repeat at several stations.
Quadrat Count all plants inside a fixed frame 1 m × 1 m frame
Fig 3. Quadrat. For plants or sessile species, place a fixed frame (e.g. 1 m × 1 m) and count every individual of each species inside it.
camera Camera Trap Motion-triggered photos over trap nights
Fig 4. Camera trap. For shy mammals, fix motion-triggered cameras, run them for equal trap nights per site, and count independent detections per species.
FIELD DATASHEET · Site: Pine Ridge SpeciesCountDetect Robin|||| |||seen Blue Tit|||| |heard Wren||||heard Chaffinch|||seen Nuthatch||seen Date 14/07 Observer RP 0630–0700 Clear
Fig 5. Field datasheet. One row per species: tally each individual, note how it was detected, and record date, observer, time, and weather. Sum per species, then paste the totals into the calculator.
Per-station tallies Robin S1:3 S2:2 S3:2Blue Tit S1:2 S2:2 S3:2 Wren S1:1 S2:2 S3:1Chaffinch S1:1 S2:1 S3:1 Nuthatch S1:1 S2:0 S3:1 add across stations → Paste into tool 7, 6, 4, 3, 2 1-D = 0.76 ✓ From datasheet to result Sum species totals, then enter
Fig 6. Sum, then enter. Add each species across all stations to get one total per species, then paste the comma-separated totals into the calculator to get Simpson's 1-D.
  1. Define your question first. Decide what you are comparing (site vs site, season vs season, before vs after) before you collect a single count.
  2. Pick one taxonomic group and stick to it. Count only birds, only trees, or only mammals. Mixing groups makes the index meaningless.
  3. Choose a standard sampling method and use it everywhere: point counts, line transects, quadrats, camera traps, pitfall traps, or mist nets.
  4. Fix your sampling effort. Keep the same transect length, plot size, count duration, or number of trap nights at every site.
  5. Set a minimum sample. Aim for at least 30 individuals and several species; small samples give unstable Simpson's values.
  6. Use replicates. Place at least 3 stations, plots, or transects per site so you can estimate variation.
  7. Mark station locations with GPS so the exact spot can be resurveyed in future years.
  8. Record every individual, not just species presence. Simpson's index needs abundance counts, not a checklist.
  9. Count during a consistent time window and avoid rain, high wind, or extreme heat.
  10. Avoid double-counting. Move in one direction, note flying birds separately, and use a set detection radius.
  11. Log detection method (seen, heard, trapped, photographed) for later corrections.
  12. Identify to species where possible. If you cannot, use a consistent morphospecies label and keep it identical across sheets.
  13. Use a prepared datasheet. Columns: Date, Site, Station, Observer, Species, Count, Detection type, Notes.
  14. Record effort metadata on every sheet: start and end time, weather, observer name, and equipment used.
  15. Photograph or voucher tricky species to settle identification disputes back at the desk.
  16. Enter counts as whole numbers, not percentages or densities.
  17. Back up your data the same day by photographing paper sheets or typing counts into a spreadsheet.
  18. Keep a field notebook for disturbance, human activity, unexpected species, or equipment problems.
  19. Sum counts per species before entry, then paste the per-species totals into this calculator (for example 52, 48, 55, 61, 47).
  20. Never invent or round zeros away. A species absent from the sample is simply not listed.

🎯 14. When to Use This Tool

14.1 Quick decision checklist

  • ✅ You have counts of individuals per species, not just a species list.
  • ✅ Your question is about dominance: is one species taking over?
  • ✅ Your sample is modest in size. Simpson's index is more stable than Shannon's at low N.
  • ✅ You need a number between 0 and 1 that a non-specialist audience can read.
  • ❌ Rare species are the point of your study. Simpson's index barely notices them.
  • ❌ You only know which species were present. Use species richness instead.
  • ❌ Effort differs a lot between sites and cannot be standardised. Rarefy first.

14.2 Four real situations

SituationWhat you doWhat 1 - D tells you
Tracking an invasive plant in a grasslandSame quadrats each year, counts per speciesWhether one species is squeezing out the rest, the exact thing Simpson detects best
Water quality monitoring on a streamKick samples above and below a dischargeWhether pollution-tolerant species now dominate downstream
Reporting a small pilot survey60 individuals, 8 species, one seasonA stable estimate where the Shannon index would still be climbing with effort
Explaining biodiversity to a council or landownerAny standardised countA plain probability: the chance two animals picked at random are different species

14.3 Sampling design guidance

  1. Fix effort first. Decide trap nights, transect length, or quadrat count before you start, and keep it identical at every site.
  2. Replicate within sites. Three or more stations per site let you report a mean with a standard error rather than a bare number.
  3. Randomise or stratify placement. Placing stations where you expect wildlife inflates the common species and drags D upward.
  4. Hold the season constant. Dominance swings with breeding, migration, and flowering. Compare like with like.
  5. Use the finite sample form for small N. Below about 100 individuals it is the more honest estimate.

14.4 Which metric next

  • Want rare species to count? Go to the Shannon Wiener index.
  • Want a fair ratio, for example "twice as diverse"? Report 1/D, which is Hill number q = 2.
  • Only care about the single commonest species? Use Berger-Parker.
  • Sample sizes differ? Use rarefaction or Margalef's index.
  • Comparing species turnover between sites? Use beta diversity, not Simpson's index.

🛠️ 15. Troubleshooting and Common Errors

Ten problems users hit with Simpson's diversity index, and how to fix each one.

My value is 0.32 but the paper I am copying reports 0.68

Cause: you calculated D, they reported 1 - D. Fix: both describe the same community. Subtract from 1, and always state which form you are reporting in your methods. This is by far the most common Simpson's index error.

My 1 - D is 0

Cause: only one species has a count above zero. Fix: nothing to fix. A single-species sample genuinely has zero diversity. Check that no species were lost during data entry.

My value is above 1

Cause: percentages were used with the finite sample formula, or the proportions do not add to 1.00. Fix: D, 1 - D and every proportion must sit between 0 and 1. Go back to raw counts.

1/D is larger than my number of species

Cause: a calculation error. Fix: 1/D can never exceed S. Check that your proportion column sums to exactly 1.00 and that no row is a total row.

Adding ten rare species barely changed my result

Cause: that is how Simpson's index works. Squaring the proportions crushes small values: 0.01 squared is 0.0001. Fix: if rare species matter to your question, report species richness or the Shannon index alongside.

The finite sample form gives a different answer

Cause: expected. The proportion form assumes an infinite population; the n(n-1) form corrects for a finite sample. Fix: use the finite form when N is below about 100, and say which you used.

R's vegan does not match my spreadsheet

Cause: vegan::diversity(x, "simpson") returns 1 - D, and it uses the proportion form. Fix: compare like with like. For D, use 1 - vegan::diversity(x, "simpson").

The tool says invalid input

Cause: text, symbols, or negative numbers in the count list. Fix: keep only whole numbers separated by commas, spaces, tabs, or new lines. Replace notes such as "none seen" with 0.

Two sites have the same 1 - D but look different

Cause: richness and evenness can trade off, and Simpson's index compresses at the top of its range. Fix: report S, 1/D and 1 - D together. The reciprocal form separates sites that 1 - D flattens.

Charts do not appear after I click Run

Cause: the Chart.js library was blocked by a network filter or an offline device. Fix: reload the page on a normal connection. The numbers and the results table still work without charts.

⚠️ 16. Assumptions and Limitations

Simpson's diversity index is robust and easy to explain, but it answers a narrower question than people assume. These are the conditions it relies on, and the things it cannot tell you.

16.1 What the index assumes

  • Individuals were sampled at random from the community.
  • All species had a fair chance of being detected. Shy or nocturnal species break this.
  • Every individual was identified to the same taxonomic level.
  • Counts are of individuals, not biomass, cover, or percentages.
  • Each record is independent. Ten photographs of one animal in one minute is one detection.
  • For the finite sample form, counts are whole numbers and N is the true total.

16.2 What it cannot tell you

  • Which species are present. Two sites with no species in common can score the same 1 - D.
  • Anything much about rare species. Squaring the proportions all but erases them.
  • Whether the species are native, threatened, or invasive. The index treats all species as equal.
  • Whether a difference between sites is statistically significant. That needs replicates and a test.
  • Population size or trend. Use density, occupancy, or abundance estimates for that.

16.3 The four limitations reviewers raise most

LimitationWhy it mattersWhat to do
Compression at the topRich sites all cluster between 0.90 and 0.99, hiding real differencesReport 1/D as well, which spreads them out
Three forms, one nameReaders cannot tell D from 1 - D from 1/D without being toldName the form explicitly in the methods and in every table caption
Blind to rare speciesLosing half your rare species may not move the index at allPair it with richness or the Shannon index
Detection biasCryptic species are systematically missed, inflating apparent dominanceUse occupancy models, or state the bias openly

What to conclude: report 1 - D as one line of evidence, with richness, 1/D, effort, and a species list beside it. On its own it is a summary, not a conclusion.

🔍 17. Conclusion

▶ Run the analysis above to generate a personalised conclusion for your dataset.

❓ 18. Frequently Asked Questions

What is Simpson's diversity index and when should I use it?

Simpson's diversity index measures the chance that two individuals picked at random from a sample belong to different species. Use it when your question is about dominance, when your sample is small, or when you need a number between 0 and 1 that a non-specialist can understand.

How do you calculate Simpson's diversity index step by step?

Divide each species count by the total to get a proportion p. Square each proportion. Add the squares together to get dominance D. Subtract from 1 for the diversity form. With counts 45, 30, 15, 7, 3 the squares sum to 0.321, so 1 - D = 0.679.

What is the difference between D, 1-D and 1/D?

D is dominance, the chance two individuals match, and higher means less diverse. 1 - D is Simpson's index of diversity, and higher means more diverse. 1/D is the reciprocal index, the effective number of common species. For the example above they are 0.321, 0.679 and 3.12.

What is the Excel formula for Simpson's diversity index?

Use =1-SUMSQ(B2:B6)/SUM(B2:B6)^2 to get 1 - D straight from raw counts in one cell. For the finite sample form use =SUMPRODUCT(B2:B6,B2:B6-1)/(B7*(B7-1)) where B7 holds the total. SUMSQ is the function most tutorials miss.

How do I calculate Simpson's index in R?

Base R needs one line: D <- sum(p^2), where p is counts divided by their total. The package version is vegan::diversity(counts, index = "simpson"), which returns 1 - D, not D. Use index = "invsimpson" for the reciprocal form.

Which Python package calculates Simpson's diversity index?

scikit-bio has skbio.diversity.alpha.simpson(), which returns 1 - D from integer counts. SciPy has no diversity function, so most people use NumPy directly: 1 - np.sum((c/c.sum())**2). Do not confuse it with scipy.integrate.simpson, which is numerical integration.

What is a good Simpson's diversity index value?

There is no universal cutoff. Values of 1 - D above 0.85 are high, 0.70 to 0.85 is healthy for most temperate systems, 0.50 to 0.70 is moderate, and below 0.50 suggests strong dominance. Always compare against a reference site sampled the same way.

What data do I need to calculate Simpson's diversity index?

You need the count of individuals for each species in your sample. Paste them comma separated, upload a CSV file, or type them into the grid. Whole numbers are required for the finite sample form. Aim for at least 30 individuals.

What does a high Simpson's index mean?

It depends on which form. A high 1 - D means high diversity, with individuals spread across many species. A high D means the opposite, with one or two species dominating. This is why naming the form in your methods matters more for Simpson than for any other index.

Simpson's index vs Shannon index, which should I use?

Simpson weights common species and is stable at small sample sizes. Shannon weights rare species and is more sensitive to richness. Report both when you can. If your question is about one species taking over, Simpson answers it better.

Why does Simpson's index ignore rare species?

Because it squares the proportions. A species holding 45 percent of individuals contributes 0.2025 to D, while one holding 1 percent contributes 0.0001. Squaring crushes small values, which is a design feature, not a flaw.

When should I use the finite sample formula?

Use D = Σ n(n-1) / [N(N-1)] when your total sample is below roughly 100 individuals. It corrects for the fact that you are sampling without replacement. Above N = 200 the two forms agree to three decimals.

Does sample size affect Simpson's diversity index?

Less than it affects the Shannon index. Because it depends on the common species, Simpson's index stabilises after relatively few individuals. It still rises slightly with effort, so standardise effort across sites before comparing.

Can I compare Simpson's index between sites or seasons?

Yes, when effort, protocol, taxonomic group and season are comparable. Report 1/D as well as 1 - D, because 1 - D compresses at the top of its range and can hide real differences between species-rich sites.

How do I report Simpson's diversity index in a paper?

Name the form, give the value, and add richness and sample size, for example: Simpson's index of diversity 1 - D = 0.679 (S = 5 species, N = 100 individuals). State whether you used the proportion form or the finite sample form.

What are the assumptions and limitations of Simpson's index?

It assumes a random, representative sample with standardised effort and correct identification. It cannot tell you which species are present, whether they are native or threatened, or anything useful about rare species.

What is the Gini-Simpson index?

It is another name for 1 - D, Simpson's index of diversity. The same quantity appears in economics as the Gini impurity and in machine learning as a splitting criterion, which is why the name travels between fields.

What is the inverse Simpson index used for?

1/D gives the effective number of common species, so it behaves like a species count. A site with 1/D of 10 really is twice as diverse as one with 5, which is not true of 1 - D. It is also Hill number q = 2.

Why does my Simpson's index not match my supervisor's?

Almost always because one of you reported D and the other reported 1 - D. The second most common cause is the finite sample correction. Compare the raw counts and the formula used before assuming either value is wrong.

Is this Simpson's diversity index calculator free and accurate?

Yes. It is completely free, runs entirely in your browser, stores no data, and uses the published Simpson formulas in all three forms. Results are cross-checked against R's vegan package and Python, and match to three decimal places.

📝 19. Cite This Tool

Use one of these three formats when you mention this Simpson's diversity index calculator in a thesis, report, or paper.

APA 7th edition

StatsUnlock. (2026). Simpson's diversity index calculator: Free online ecology tool [Computer software]. https://statsunlock.com/simpsons-diversity-index-calculator/

BibTeX

@misc{statsunlock_simpson_2026,
  title        = {Simpson's Diversity Index Calculator: Free Online Ecology Tool},
  author       = {{StatsUnlock}},
  year         = {2026},
  howpublished = {\url{https://statsunlock.com/simpsons-diversity-index-calculator/}},
  note         = {Accessed: 2026-09-15}
}

Methods paragraph

Species diversity was summarised using Simpson's index (Simpson, 1949; Magurran, 2004). Dominance was calculated as D = Σ p², where p is the proportion of individuals belonging to each species, and results are reported as Simpson's index of diversity (1 - D) together with the reciprocal form (1/D). All indices were computed with the StatsUnlock Simpson's diversity index calculator (https://statsunlock.com/simpsons-diversity-index-calculator/) and verified in R with vegan::diversity().

🔗 20. Related Tools

Free ecology and wildlife calculators that pair well with Simpson's diversity index.

Shannon Wiener Index Calculator

The companion index. It weights rare species, where Simpson weights common ones. Report both.

Evenness Index Calculator (Pielou's J')

Separates evenness from richness when two sites share the same 1 - D.

Hill Numbers Calculator

Puts 1/D on a common scale with richness and the Shannon index. 1/D is Hill q = 2.

Relative Abundance Calculator

Turns raw counts into the proportions the Simpson formula squares.

Fisher's Alpha Diversity Calculator

A richness measure that is less sensitive to sample size.

Camera Trap Analysis Calculator

Turns photo detections and trap nights into the standardised counts this tool needs.

📖 21. Glossary of Terms

Abundance The number of individuals of a species recorded in a sample.
Alpha diversity Diversity within a single site or sample. Simpson's index is an alpha diversity measure.
Berger-Parker index The proportion of the sample made up by the single commonest species.
Beta diversity The change in species composition between sites.
Detection probability The chance a species present at a site is actually recorded during a survey.
Dominance (D) The chance that two individuals drawn at random belong to the same species.
Effective number of species 1/D, the number of equally common species that would give the same dominance.
Evenness How equally individuals are shared among species. Simpson's version is (1/D) divided by S.
Finite sample correction The n(n-1) form of D, used when the sample is small relative to the population.
Gini-Simpson index Another name for 1 - D, the Simpson index of diversity.
Hill number A family of diversity measures controlled by an order q. At q = 2 it equals 1/D.
Independent detection One record of a species separated from the next by a set time, usually 30 minutes for camera traps.
Inverse Simpson Another name for the reciprocal index, 1/D.
Proportion (p) A species count divided by the total count. All proportions in a sample add to 1.00.
Quadrat A fixed-size sampling frame, usually square, placed on the ground to count plants or slow-moving animals.
Rarefaction Standardising samples to the same number of individuals so diversity can be compared fairly.
Sampling effort The amount of survey work done: trap nights, transect kilometres, point counts, or quadrats.
Shannon Wiener index (H') A diversity index that weights rare species more heavily than Simpson's index does.
Simpson's index of diversity 1 - D, the chance two random individuals are different species.
Simpson's reciprocal index 1/D, the effective number of common species, ranging from 1 to S.
Species accumulation curve A plot of species found against effort. When it flattens, sampling is adequate.
Species richness (S) The number of different species in a sample, with no weighting by abundance.
Trap night One camera trap active for 24 hours. The standard unit of camera trap effort.
Transect A fixed line along which observations are recorded at a steady speed and effort.
📚 22. References (20 Peer-Reviewed Sources)

The following references support the ecological methods used in this Simpson's diversity index calculator, covering biodiversity measurement, species diversity, and best practices in ecological sampling and wildlife analysis.

  1. Simpson, E. H. (1949). Measurement of diversity. Nature, 163, 688. doi.org/10.1038/163688a0
  2. Shannon, C. E., & Weaver, W. (1949). The mathematical theory of communication. University of Illinois Press.
  3. Magurran, A. E. (2004). Measuring biological diversity. Blackwell Publishing.
  4. Krebs, C. J. (1999). Ecological methodology (2nd ed.). Benjamin Cummings.
  5. Hill, M. O. (1973). Diversity and evenness: A unifying notation and its consequences. Ecology, 54(2), 427–432. doi.org/10.2307/1934352
  6. Jost, L. (2006). Entropy and diversity. Oikos, 113(2), 363–375. doi.org/10.1111/j.2006.0030-1299.14714.x
  7. Lande, R. (1996). Statistics and partitioning of species diversity. Oikos, 76(1), 5–13. doi.org/10.2307/3545743
  8. Chao, A., & Jost, L. (2012). Coverage-based rarefaction and extrapolation. Ecology, 93(12), 2533–2547. doi.org/10.1890/11-1952.1
  9. Oksanen, J., et al. (2022). vegan: Community ecology package. R package v2.6-4. CRAN.R-project.org/package=vegan
  10. Bibby, C. J., Burgess, N. D., Hill, D. A., & Mustoe, S. H. (2000). Bird census techniques (2nd ed.). Academic Press.
  11. Gotelli, N. J., & Colwell, R. K. (2001). Quantifying biodiversity: Procedures and pitfalls. Ecology Letters, 4(4), 379–391. doi.org/10.1046/j.1461-0248.2001.00230.x
  12. Colwell, R. K. (2013). EstimateS: Statistical estimation of species richness (v9). purl.oclc.org/estimates
  13. Whittaker, R. H. (1972). Evolution and measurement of species diversity. Taxon, 21(2/3), 213–251. doi.org/10.2307/1218190
  14. R Core Team. (2024). R: A language and environment for statistical computing. R-project.org
  15. Ahumada, J. A., et al. (2011). Community structure and diversity of tropical forest mammals. Phil. Trans. R. Soc. B, 366(1578), 2703–2711. doi.org/10.1098/rstb.2011.0115
  16. Chao, A., et al. (2014). Rarefaction and extrapolation with Hill numbers. Ecological Monographs, 84(1), 45–67. doi.org/10.1890/13-0133.1
  17. Niedballa, J., et al. (2016). camtrapR: An R package for camera trap data. Methods Ecol. Evol., 7(12), 1457–1462. doi.org/10.1111/2041-210X.12600
  18. Hsieh, T. C., Ma, K. H., & Chao, A. (2016). iNEXT: rarefaction and extrapolation of Hill numbers. Methods Ecol. Evol., 7(12), 1451–1456. doi.org/10.1111/2041-210X.12613
  19. Roswell, M., Dushoff, J., & Winfree, R. (2021). A conceptual guide to measuring species diversity. Oikos, 130(3), 321–338. doi.org/10.1111/oik.07202
  20. Peet, R. K. (1974). The measurement of species diversity. Annual Review of Ecology and Systematics, 5, 285–307. doi.org/10.1146/annurev.es.05.110174.001441

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